Manzanitas (Arctostaphylos, Ericaceae family) have proliferated in California's Mediterranean climate in multiple ways. Tom will introduce you to the diversity of this group, and then focus on two stories. The first story is where manzanitas come from, so we will examine the history of a small subfamily, the Arbutoideae, of the blueberry/rhododendron/heath family (Ericaceae). Most of the genera in this subfamily occur in California, but only two are in northern California. This history leads into the second story, how manzanitas have captured mutualists, rodents such as chipmunks and chaparral mice, that bury fruits that help them survive our fires.